Academic appointments

Vanderbilt University Medical Center, 2000-present. Assistant Professor, Division of Surgical Oncology; 2004-present: Assistant Professor, Department of Cell and Developmental Biology.

 Professional Associations

American Association for Cancer Research

Society for Developmental Biology

American Society for Cell Biology

American Pancreatic Association


Vanderbilt-Ingram Cancer Center

Vanderbilt Digestive Disease Research Center

Vanderbilt Diabetes Research and Training Center

Vanderbilt Program in Developmental Biology



The following timeline graph is generated from all co-authored publications.

Featured publications are shown below:

  1. Prox1-Heterozygosis Sensitizes the Pancreas to Oncogenic Kras-Induced Neoplastic Transformation. Drosos Y, Neale G, Ye J, Paul L, Kuliyev E, Maitra A, Means AL, Washington MK, Rehg J, Finkelstein DB, Sosa-Pineda B (2016) Neoplasia 18(3): 172-84
    › Primary publication · 26992918 (PubMed) · PMC4796801 (PubMed Central)
  2. Expression of CD24, a Stem Cell Marker, in Pancreatic and Small Intestinal Neuroendocrine Tumors. Salaria S, Means A, Revetta F, Idrees K, Liu E, Shi C (2015) Am J Clin Pathol 144(4): 642-8
    › Primary publication · 26386086 (PubMed) · PMC4576728 (PubMed Central)
  3. Loss of HNF6 expression correlates with human pancreatic cancer progression. Pekala KR, Ma X, Kropp PA, Petersen CP, Hudgens CW, Chung CH, Shi C, Merchant NB, Maitra A, Means AL, Gannon MA (2014) Lab Invest 94(5): 517-27
    › Primary publication · 24638272 (PubMed) · PMC4068339 (PubMed Central)
  4. Fibrogenesis in pancreatic cancer is a dynamic process regulated by macrophage-stellate cell interaction. Shi C, Washington MK, Chaturvedi R, Drosos Y, Revetta FL, Weaver CJ, Buzhardt E, Yull FE, Blackwell TS, Sosa-Pineda B, Whitehead RH, Beauchamp RD, Wilson KT, Means AL (2014) Lab Invest 94(4): 409-21
    › Primary publication · 24535260 (PubMed) · PMC3992484 (PubMed Central)
  5. Heparin-binding epidermal growth factor-like growth factor eliminates constraints on activated Kras to promote rapid onset of pancreatic neoplasia. Ray KC, Moss ME, Franklin JL, Weaver CJ, Higginbotham J, Song Y, Revetta FL, Blaine SA, Bridges LR, Guess KE, Coffey RJ, Crawford HC, Washington MK, Means AL (2014) Oncogene 33(7): 823-31
    › Primary publication · 23376846 (PubMed) · PMC3929321 (PubMed Central)
  6. Pancreatic stellate cells: small cells with a big role in tissue homeostasis. Means AL (2013) Lab Invest 93(1): 4-7
    › Primary publication · 23269285 (PubMed)
  7. Epithelial tissues have varying degrees of susceptibility to Kras(G12D)-initiated tumorigenesis in a mouse model. Ray KC, Bell KM, Yan J, Gu G, Chung CH, Washington MK, Means AL (2011) PLoS One 6(2): e16786
    › Primary publication · 21311774 (PubMed) · PMC3032792 (PubMed Central)
  8. Adult pancreatic acinar cells give rise to ducts but not endocrine cells in response to growth factor signaling. Blaine SA, Ray KC, Anunobi R, Gannon MA, Washington MK, Means AL (2010) Development 137(14): 2289-96
    › Primary publication · 20534672 (PubMed) · PMC2889602 (PubMed Central)
  9. Pancreatic ductal morphogenesis and the Pdx1 homeodomain transcription factor. Wescott MP, Rovira M, Reichert M, von Burstin J, Means A, Leach SD, Rustgi AK (2009) Mol Biol Cell 20(22): 4838-44
    › Primary publication · 19793922 (PubMed) · PMC2777112 (PubMed Central)
  10. Transmembrane and soluble isoforms of heparin-binding epidermal growth factor-like growth factor regulate distinct processes in the pancreas. Ray KC, Blaine SA, Washington MK, Braun AH, Singh AB, Harris RC, Harding PA, Coffey RJ, Means AL (2009) Gastroenterology 137(5): 1785-94
    › Primary publication · 19689925 (PubMed) · PMC2767440 (PubMed Central)
  11. Epidermal growth factor receptor regulates pancreatic fibrosis. Blaine SA, Ray KC, Branch KM, Robinson PS, Whitehead RH, Means AL (2009) Am J Physiol Gastrointest Liver Physiol 297(3): G434-41
    › Primary publication · 19608732 (PubMed) · PMC2739824 (PubMed Central)
  12. A CK19(CreERT) knockin mouse line allows for conditional DNA recombination in epithelial cells in multiple endodermal organs. Means AL, Xu Y, Zhao A, Ray KC, Gu G (2008) Genesis 46(6): 318-23
    › Primary publication · 18543299 (PubMed) · PMC3735352 (PubMed Central)
  13. Pancreatic epithelial plasticity mediated by acinar cell transdifferentiation and generation of nestin-positive intermediates. Means AL, Meszoely IM, Suzuki K, Miyamoto Y, Rustgi AK, Coffey RJ, Wright CV, Stoffers DA, Leach SD (2005) Development 132(16): 3767-76
    › Primary publication · 16020518 (PubMed)
  14. The islet beta cell-enriched RIPE3b1/Maf transcription factor regulates pdx-1 expression. Samaras SE, Zhao L, Means A, Henderson E, Matsuoka TA, Stein R (2003) J Biol Chem 278(14): 12263-70
    › Primary publication · 12551916 (PubMed)